Plants terrestrial or on rock [epiphytic]. Stems creeping to erect, scaly at apex. Leaves monomorphic or somewhat dimorphic [dimorphic]. Petiole in cross section with 2 crescent-shaped vascular bundles at base. Blade pinnate to pinnate-pinnatifid, rarely more than 2-pinnate [simple]; rachis grooved adaxially or not, grooves not continuous with grooves of next order. Veins free or anastomosing, running to margin, areoles with or without included free veinlets. Indument of transparent, needlelike, hooked, septate, or stellate hairs, or rarely hairs lacking. Sori inframedial to supramedial, occasionally nearly marginal, round or oblong, rarely elongate along veins; indusia reniform or sometimes absent. Spores bilateral, monolete [rarely globose-tetrahedral and trilete], usually with a prominent, crested, echinate, or reticulate perispore. Gametophytes green, cordate, usually hairy or glandular; antheridia 3-celled.
Plants terrestrial or on rocks. Rhizomes stout, dictyostele radially symmetrical, branched or not, erect, ascending, or long creeping, with scales at apices; scales basiflexed, lanceolate or nearly ovate, brown, thick, luminae elongate, usually with grayish white short setae on dorsal side or ciliate along margins. Fronds clustered, approximate, or remote; stipes slender, stramineous, not articulate, with 2 crescent-shaped vascular bundles at base, usually scaly at bases, distally ± with grayish white unicellular acicular hairs, rarely with multicellular long hairs or stellate hairs. Fronds monomorphic, rarely subdimorphic, oblong-lanceolate or oblanceolate, sometimes ovate or ovate-triangular, usually pinnate-pinnatifid, sometimes 3- or 4-pinnate-pinnatifid, rarely 1-pinnate; pinnae symmetrical at bases; costae grooved adaxially but grooves not confluent with rachial grooves, or raised and with dense grayish acicular hairs, with expanded tuberculate aerophores at bases of pinnae. Laminae herbaceous or papery, sometimes somewhat leathery, green or dark brown-green when dry, both sides (particularly rachises, costae, and main veins adaxially) with grayish white unicellular acicular hairs, rarely glabrous, usually with orange or reddish orange, stalked or sessile spherical or club-shaped glands, occasionally small scaly along rachises and costae abaxially. Sori orbicular, oblong, or shortly linear, dorsifixed on veins, indusiate or exindusiate; indusia orbicular-reniform, fixed by deep notch, most ± hairy, persistent or hidden in sori, caducous, or not concentrated into sori but scattered along reticulate veins and exindusiate. Sporangia long stalked, usually with hairs or glandular hairs below annuli and at distal end of sporangial stalks. Spores bilateral, rarely tetrahedral, tuberculate, echinate, granular, or usually with a winged perispore. Prothalli green, cordate or narrowly cordate, usually with broad wings, symmetrical, usually with hairs or glands. x = 27-36 (lacking 28).
|Marsh Fern Family|