Plants terrestrial, epilithic, or epiphytic. Rhizome long, creeping, erect, or scandent; scales blackish brown, thick, spreading or appressed, imbricate, peltate at base, margin often long ciliate. Fronds distant or clustered; stipe articulate to raised phyllopodia; lamina simple, entire, lanceolate to linear-lanceolate, herbaceous, papery, or leathery, margin cartilaginous, glabrous or pubescent; costa prominent, raised adaxially, often with small scales on abaxial surface; veins simple or forked, free. Sori in a single often irregular row on either side of costa; indusia persistent, red-brown, reniform or orbicular-reniform, membranous or papery. Spores monolete, with broad, winglike, echinulate folds, cristate or echinate to erose.