Plants medium-sized or large, usually in forests near or beside streams. Rhizomes decumbent to erect, thick. Lamina 1-pinnate, imparipinnate, with pinnae entire, glabrous; veins free on either side of midrib, fully anastomosing toward margins and forming 2-4 rows of areoles, without included free veinlets. Sori elongate along long veins close to midrib, covered by a similarly elongate, membranous indusium; indusia usually 3-8 mm, sometimes up to 1-2 cm, arched, sausage-shaped. Spores with winglike fimbriate and echinate folds, or low perforate plain folds. x = 41, 40 or 31.