Athyriaceae Alston
  • Taxon 5: 25. 1956. (25 Mar 1956)

This taxon is accepted by World Flora Online consortium
Notes: More details could be found in The Plant List v.1.1.

General Description

Plants terrestrial or epilithic, small to medium-sized, sometimes large. Rhizome long or shortly creeping, ascending or erect, rarely short treelike, dictyostelic, scaly; scales entire or toothed, tooth consisting of 2 adjacent upturned ends of marginal cells, basifixed, rarely pseudopeltate. Stipe supplied by 2 vascular strands with xylem seahorse-shaped in cross section and upward united into U-shaped strand, scaly, also hairy or glabrate, hairs unicellular or multicellular, glandular or eglandular. Rachis, costae, and costules grooved adaxially, grooves of costae and costules confluent to those of lower order frond axes or interrupted by ridges between grooves, scaly, hairy, or glabrate; hairs unicellular or multicellular, glandular or eglandular; spines continued from ridges, or short fleshy projections in grooves, at base of costae, costules, and midribs adaxially, or spines and projections absent. Lamina variously dissected, simple to 3-pinnate, pinnatifid or with terminal pinna similar to lateral pinnae, apex narrowed. Veins free or anastomosing with polygonal areoles or with successive veins from adjacent midribs united to excurrent veins between 2 rows of rhomboid areoles. Sori various, linear, J-shaped, horseshoe-shaped, orbicular-reniform, or orbicular, indusiate or exindusiate; indusia inferior entirely or proximally to receptacle, or lateral, various in shape, sack-shaped, cup-shaped, saucer-shaped, platelike, elongate, J-shaped, horseshoe-shaped, or orbicular-reniform. Spores bilateral, ellipsoid, covered by perispore.

Five genera and ca. 600 species: worldwide from tropical to frigid zones of both hemispheres; five genera and 278 species (135 endemic) in China.

Of the genera recognized in FRPS (3(2): 32-504. 1999), Rhachidosorus and Diplaziopsis were recently segregated as families Rhachidosoraceae and Diplaziopsidaceae, respectively. Athyriaceae s.s. or subfamily Athyrioideae comprise Anisocampium (including Kuniwatsukia), Athyrium (including Pseudocystopteris), Cornopteris (including Neoathyrium), Deparia (including Athyriopsis, Dictyodroma, Dryoathyrium, and Lunathyrium), and Diplazium (including Allantodia, Callipteris, and Monomelangium). Subfamily Cystopteridoideae was raised to family Cystopteridaceae.

Wang Zhong-ren, Chu Wei-ming, He Zhao-rong & Hsieh Yin-tang. 1999. Athyriaceae (excluding Acystopteris, Cystoathyrium, Cystopteris, Diplaziopsis, Gymnocarpium, and Rhachidosorus). In: Chu Wei-ming, ed., Fl. Reipubl. Popularis Sin. 3(2): 32-511.


"Rhizomes terrestrial, usually short, stout and suberect; scales not clathrate, with entire or toothed margins"


  • 1
  • 2 Taxon 5: 25. 1956. (25 Mar 1956)

 Information From

Flora Of China @
'Flora of China @ eFloras (2008). Published on the Internet [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
  • A All Rights Reserved
Plants Of the World Online Portal
  • B
World Flora Online Data. 2017.
  • C CC0 1.0 Universal (CC0 1.0).
World Flora Online consortium
World Flora Online Data. 2017.
  • D CC0 1.0 Universal (CC0 1.0).