Shrubs, climbing, sometimes erect or small trees, indument with stellate hairs. Inflorescences terminal, axillary, extra-axillary, leaf-opposed, or rarely on older branches; flowers mostly solitary or paired, rarely in short racemes or few-flowered cymes. Torus depressed, pubescent or tomentose. Sepals 3, valvate, usually connate at base. Petals 6, in 2 whorls, sometimes connate at very base, each whorl imbricate. Stamens many; anther locules often oblong or linear; connectives apically truncate to rounded. Carpels many, rarely few, linear-oblong; ovules many per carpel or rarely 2 or 3, in 1 or 2 series; styles outside often pilose; stigmas usually apically 2-cleft, involute. Fruit apocarpous; monocarps mostly ± stipitate, oblong, ovoid, or subglobose, fleshy and berrylike. Seeds several per monocarp or 1 by abortion, subhorizontal, with or without a short aril.
Seeds horizontal or oblique, with or without a small aril.
Petals 6, imbricate, in 2 whorls of 3, equal or those of the outer whorl slightly larger, free or scarcely joined at the base
Sepals 3, valvate, often ± joined at the base, or calyx cupular in bud and remaining ± unlobed in anthesis or splitting into 3 often irregular lobes
Flowers usually hermaphrodite, solitary or in few-flowered cymes, these terminal, axillary, extra-axillary or leaf-opposed, more rarely borne on the old wood, sessile or stalked; bracteoles 2, respectively basal and median or at the base of the calyx, often deciduous
Climbing, straggling or erect shrubs or small (less often quite large) trees, quite often covered with stellate and sometimes also simple hairs
Monocarps indehiscent, globose to cylindrical, stipitate or subsessile, 1–many-seeded, sometimes grooved between the seeds, dry or sometimes fleshy when ripe, often sweet and edible
Carpels mostly numerous, free, linear or cylindric, with numerous biseriate ovules (rarely with only 1–3); style absent or scarcely developed; stigma horseshoe-shaped or funnel-shaped with a slit down the inner side
Stamens numerous, the outer sometimes sterile and leaf-like, linear or obconic; anther-cells lateral or extrorse; connective truncate or prolonged into a tapering appendage; filaments very short or absent
Seeds horizontal or oblique; aril small, carunculoid.
Carpels ?, free, linear or cylindric, with numerous ovules in 2 rows, rarely with 1–3 ovules; style very short or absent; stigma horse-shoe-shaped or funnel-shaped with a slit down the inner side.
Ripe carpels indehiscent, dry or succulent, often sweet and edible, globular to cylindric, usually on median or lateral stipes, many- to few-seeded.
Petals 6, in two whorls, both imbricate, equal or the outer whorl slightly longer, expanding and spreading at anthesis, free or rarely connate at the base.
Stamens ?, the outer sometimes sterile and foliar, linear or obconic, with thecae lateral or extrorse and connective prolongations thin, tapering, rounded or truncate, rarely capitate; filaments very short or absent.
Bracteoles 2, respectively basal and median or at the base of the calyx, often caducous.
Sepals 3, valvate, often ± connate.
Flowers usually bisexual, solitary or in few-flowered cymes, terminal or extra-axillary (often leaf-opposed), rarely on old branches, sessile or pedicellate.