Plants medium-sized to large, in loose tufts or extensive mats, deep green, light green, or whitish, brownish or paler green with age. Stems creeping, terete- to subcomplanate-foliate, not julaceous, irregularly branched, branches terete-, complanate-, or subcomplanate-foliate; central strand present; pseudoparaphyllia acute or acuminate; axillary hairs of 3-7 cells, . Stem leaves erectopatent or erect, gradually reflexed from erect base, imbricate to somewhat spaced, broadly ovate or suborbicular to ovate-lanceolate, flat to slightly concave, not or slightly plicate; base with decurrency absent after leaf is detached; margins serrate to serrulate; apex acuminate or ± broadly acute; costa to 40-85% leaf length, moderate to somewhat stout, terminal abaxial spine present or absent; alar cells poorly differentiated, or short-rectangular, slightly enlarged, ; laminal cells linear, walls moderately thick; basal cells somewhat shorter. Branch leaves similar, smaller, sometimes narrower. Sexual condition autoicous; perichaetial leaves abruptly contracted, acumen straight to reflexed. Seta red-brown, smooth. Capsule inclined to horizontal, red-brown to brown, oblong-cylindric, weakly curved; annulus separating; operculum rostrate; peristome xerocastique, perfect. Calyptra naked. Spores 9-16 µm.