Herbs perennial. Rootstock short, crown usually bearing remnant leaf sheaths. Stem finely striate, dichotomously branched above. Leaves petiolate; petioles sheathing. Umbels loosely compound, terminal and lateral; bracts numerous or absent; rays numerous or few; bracteoles numerous, rarely few or absent. Calyx teeth short or obsolete. Petals usually white, occasionally pinkish or purplish, rarely pale yellow, orbicular to obovate, with a narrow inflexed apex. Stylopodium low-conic. Fruit ellipsoid, oblong to suborbicular, dorsally compressed; dorsal ribs filiform, slightly prominent, lateral ribs thickened and narrowly winged, wings closely appressed to one another; vittae 1 to several in each furrow, 2 to several on commissure. Seed face plane or slightly concave. Carpophore bipartite.
"Fr strongly flattened dorsally, the lateral ribs evidently winged, the others wingless; oil-tubes 1–3 in the intervals, 2 or more on the commissure; carpophore bifid to the base; umbels compound; pet white or yellow; stylopodium conic; perennials, usually coarse and erect from a taproot or stout rhizome, with pinnately or ternately compound or dissected lvs, often with well defined lfls. 100+, Old World."
Leaves variously pinnately or ternately divided into filiform to broad and jaggedly toothed segments, nervation pinnate, very rarely parallel-nerved and then with obscure secondary venation
Endosperm flat or shallowly bisulcate on the commissural face, slightly to clearly sulcate beneath the vittae dorsally.
Biennial or perennial herbs, glabrous to variously pilose, rootstock various
Fruit strongly dorsally compressed, oblong to elliptic, obovate or roundish, glabrous, truncate to slightly emarginate at base and apex, the stylopodia ± apical, commissure wide; 3 dorsal ribs ± prominent, slender, 2 marginal ribs expanded into distinct, broad to rather narrow, paper-thin or thicker wings; vittae 2–8(–12) on the commissural face, when numerous then slender and variable in number and sometimes irregular; vallecular vittae l–3(–5) or rarely almost a complete ring of vittae around the endosperm; vittae usually black and conspicuous in the ripe fruit, but on the commissure sometimes concealed by overlying tissue and clear only in section; stylopodia conical; styles short; carpophore deeply bipartite
Umbels compound, few–many; involucre of 0 or few caducous to numerous persistent bracts; involucel usually conspicuous, of numerous bracteoles; flowers usually polygamous, the terminal umbels of stem and branches hermaphrodite and fertile or the inner partial umbels or their inner flowers ♂, the opposite or verticillate outer umbels entirely or in part ♂
Petals white to cream, yellow or (commonly in Africa) greenish cream, ± incised above with a large or smaller incurved acute point
Calyx-teeth absent to small but distinct
Lateral wings well developed as expansions of the marginal ribs, sometimes surrounding the stylopodium on both sides, dorsal ribs filiform.
Fruit orbicular, oblong or pyriform, very strongly dorsiventrally compressed, with a broad commissure.
Petals white, cream or yellow, sometimes flushed with pink.
Umbels compound, bracts 0 to numerous, probably sometimes caducous; bracteoles several to numerous.
Leaves pinnate or ternate, or compound forms of these, rarely finely divided.
Perennial herbs, rarely biennials or annuals.
Vittae 1–3 in each interval and 2 in the commissural face.