Plants medium-sized, in loose tufts, light green to whitish or brownish. Stems creeping to arching, loosely, occasionally more densely terete-foliate, julaceous or not, irregularly or sometimes regularly branched, branches terete-, subcomplanate-, or sometimes distinctly complanate-foliate, especially when leaves remote; central strand present; pseudoparaphyllia acute; axillary hairs of 2 or 3 cells. Stem leaves erectopatent to patent, loosely arranged to occasionally loosely imbricate, broadly ovate to ovate-lanceolate, slightly to moderately strongly concave, not plicate; base narrowly short-decurrent; margins serrate to serrulate; apex gradually tapered to acute, long- or short-acuminate; costa to 50-70% leaf length, broad throughout, terminal spine present, sometimes with additional teeth abaxially; alar cells enlarged; laminal cells elongate, walls moderately thick; basal juxtacostal cells shorter, slightly broader. Branch leaves slightly to strongly differentiated, if strongly differentiated then more elliptic, broadest at 1/3 -1/2 leaf length, twisted mid leaf; base often asymmetric; apex more shortly acuminate to acute; costa more strongly toothed distally, terminal spine more stout; . Sexual condition dioicous; perichaetial leaf acumen long, reflexed. Seta red-brown, rough. Capsule inclined to horizontal, red-brown, elongate, curved; annulus separating in fragments; operculum short-conic, narrowly long-rostrate; peristome xerocastique, perfect. Calyptra naked. Spores 10-13 µm.