Plants small to large, yellow-green, green, dark green, or blackish. Stems unbranched or irregularly branched; hyalodermis rare, central strand well developed, rarely poorly so, or absent; paraphyllia absent; rhizoids or rhizoid initials on stem or abaxial costa insertion, rarely forming tomentum, slightly to strongly branched, smooth or rarely warty-papillose; axillary hairs (well developed and many, or small, delicate, sparse), hyaline when young. Leaves recurved, squarrose, or not, straight, falcate-secund, or both, ovate, broadly ovate, oval, orbicular, ovate-lanceolate, oblong-lanceolate, or lanceolate, not plicate, to or longer than 1 mm; base commonly decurrent; margins usually plane, sometimes recurved in apex, entire or weakly denticulate or serrulate, limbidia absent; apex broadly rounded, obtuse, or acute, occasionally apiculate from obtuse or rounded apex, sometimes blunt from acute apex, ; costa usually double, short, ending before mid leaf, often single and slender or stout to 1/2-3/4 leaf length, occasionally double and long, rarely percurrent, ; alar cells undifferentiated or little differentiated, quadrate, short-rectangular, or irregular, or cells enlarged, not or slightly inflated, hyaline, yellow-brown, brown, reddish brown, or rarely bright red, , region mostly well delimited, variously small and irregular to large; medial laminal cells usually linear-flexuose, sometimes short-rhombic or fusiform, <27- 66 µm (when costa is double); distal cells sometimes prorate distally on dorsal side>; marginal cells short or very long. Sexual condition autoicous or dioicous. Capsule erect, symmetric, or inclined, ovoid to oblong-cylindric, slightly to strongly arcuate; peristome double; exostome margins almost entire to dentate; endostome cilia 1-3, absent or rudimentary to well developed, sometimes appendiculate. Spores 10-24 µm.