Cyathula Blume
  • Bijdr. Fl. Ned. Ind. 548. 1826. (24 Jan 1826)


Cite taxon page as 'WFO (2020): Cyathula Blume. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-4000010219. Accessed on: 18 Feb 2020'

General Information

Herbs or subshrubs. Stem erect or ascending. Leaves opposite, petiolate, margin entire. Flowers clustered in cymose partial inflorescences, 1-3 in each cluster, hermaphroditic, partly accompanied by sterile ones. Bracts ovate, membranous, often spiny. Tepals 5, nearly equal, membranous. Stamens 5; anther 2-loculed, oblong; filaments connate into a short cup at base, alternating with dentate or lacerate staminodes. Ovary obovoid; ovule 1, pendulous from a long funicle; style filiform, persistent; stigma capitulate. Utricles included in persistent perianth, globose, ellipsoid, or obovoid, membranous, indehiscent. Seeds oblong or ellipsoid.

  • Provided by: [F].Flora Of CHina @ efloras.org
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    CYATHULA Blume

    Hierbas o subarbustos, anuales o perennes, tallos erectos, ascendentes o escandentes, subglabros o pubescentes con tricomas simples; plantas hermafroditas. Hojas opuestas, decurrentes sobre el pecíolo, subglabras o pubescentes con tricomas simples; pecioladas. Inflorescencias cimosas parciales, terminales en el tallo y las ramas, y en las axilas de los pares de hojas caulinares superiores, cimas arregladas en estructuras espiciformes cortas, compactas o alargadas, a veces en tirsos, bráctea angostamente ovada o algo triangular, deflexa en la madurez, dorsalmente glabra, margen frecuentemente con tricomas cortos, simples, el nervio principal formando un mucrón o excurrente, bractéolas glabras, cresta ausente, las de primer orden con el nervio principal excurrente formando una cerda recta o ligeramente encorvada, las de segundo orden u órdenes superiores con el nervio principal largamente excurrente formando una cerda recta o un gancho fuerte, flores bisexuales o estériles y entonces frecuentemente rudimentarias; tépalos 5, lanceolados u oblongos, libres en la base, escariosos, margen hialino, escasa o densamente pubescentes con tricomas simples, 1–5-nervios, tépalos de las flores estériles frecuentemente modificados en una cerda con alas laterales membranáceas; estambres 5, filamentos filiformes, unidos en una cúpula poco profunda, lobos de los filamentos ausentes, anteras biloculares; pseudoestaminodios presentes, truncados, dentados o fimbriados en el ápice, más cortos que los filamentos; ovario 1-ovulado, estigma penicilado. Fruto un utrículo membranáceo, partido irregularmente por el crecimiento de la semilla, arilo ausente; flor cayendo junto con las bractéolas en la madurez.

    Género con ca 25 especies, originario del Viejo Mundo pero en la actualidad en los trópicos y subtrópicos de ambos hemisferios; 2 especies se han extendido como malezas en toda América tropical y se encuentran en Nicaragua. El nombre C. triuncinata var. managuensis Ram. Goyena de la Flora Nicaragüense es dudoso ya que la especie es endémica de Madagascar.

  • Provided by: [B].Flora de Nicaragua
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    Flowers perfect, some sterile and highly modified, bracteate and bibracteolate, subsessile in short-pedunculate glomerules. Sepals 5, discrete, hypogynous, concave, subequal, 1-3 ribbed, occasionally pungent. Stamens 5, the filaments flattened and united below into an appendiculate tube; anthers 4-locellate, introrse, medially attached. Ovary ovoid, 1-locular, 1-ovulate; ovules campylotropous on elongate flattened funicles; style 1, filiform; stigma 1, capitate. FruLit an ovoid indehiscent utricle; seeds cochleate-orbiculate, succineous. Pubescent branched ascending to prostrate occasionally frutescent herbs. Leaves opposite, entire, petiolate, strigose, deciduous. Inflorescences spicate, composed of terminal and axillary thyrses of complex glomerules containing some highly modified flowers with some of the bracts and sepals uncinate-spinescent.

  • Provided by: [E].Flora de Panama
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    Morphology

    Perianth segments 5, very shortly mucronate or some (especially the outer 2) hooked-aristate, serving with the bracteoles and modified flowers to distribute the fruit.

  • Provided by: [D].Plants Of the World Online Portal - FZ
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    Stamens 5, the filaments delicate, shortly monadelphous at the base, alternating with distinct, commonly toothed or lacerate pseudostaminodes; anthers bilocular.

  • Provided by: [D].Plants Of the World Online Portal - FZ
    • Source: [
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    Style slender, stigma capitate.

  • Provided by: [D].Plants Of the World Online Portal - FZ
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    Ovary with a single pendulous ovule.

  • Provided by: [D].Plants Of the World Online Portal - FZ
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    Annual or perennial herbs with entire, opposite leaves.

  • Provided by: [D].Plants Of the World Online Portal - FZ
    • Source: [
    • 5
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    Inflorescence terminal on the stem and branches, spiciform or capitate, bracteate, the ultimate division basically a triad of fertile flowers, the outer pair bracteolate and subtended laterally by 2 modified bracteolate flowers consisting of a number of sharply uncinately hooked (more rarely straight or glochidiate) spines or bracteoliform processes, but one or both of the outer pair sometimes absent, bracteoles also sometimes with a hooked arista; spines of the modified flowers at first small, rapidly accrescent, few to many, clustered with the clusters not or very shortly stalked.

  • Provided by: [D].Plants Of the World Online Portal - FZ
    • Source: [
    • 5
    • ]. 

    Bracts persistent, finally more or less deflexed; bracteoles and perianth falling with the fruit.

  • Provided by: [D].Plants Of the World Online Portal - FZ
    • Source: [
    • 5
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    Fruit a thin-walled utricle, irregularly ruptured by the developing seed.

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    • 5
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    Seed ovoid, slightly compressed; endosperm copious.

  • Provided by: [D].Plants Of the World Online Portal - FZ
    • Source: [
    • 5
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    Ovary with a single pendulous ovule; style slender; stigma capitate

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
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    Fruit a thin-walled capsule, irregularly ruptured by the developing seed

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
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    Perianth-segments 5, very shortly mucronate or some (especially the outer 2) hooked-aristate, serving with the bracteoles and modified flowers to distribute the fruit

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
    • ]. 

    Stamens 5, the filaments delicate, shortly monadelphous at the base, alternating with distinct, commonly toothed or lacerate pseudostaminodes; anthers bilocular

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
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    Seed ovoid, slightly compressed; endosperm copious.

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
    • ]. 

    Inflorescence terminal on the stem and branches, spiciform or capitate, bracteate, the ultimate division basically a triad of fertile flowers, the outer pair bracteolate and subtended on the outer surface by 2 modified bracteolate flowers consisting of a number of sharply uncinately hooked (rarely glochidiate or sometimes straight) spines or bracteoliform processes, but one or both of the outer pair sometimes absent, bracteoles sometimes also with a strongly hooked arista, the spines of the modified flowers at first small, rapidly accrescent, few to many, clustered, with the clusters not or very shortly stalked

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
    • ]. 

    Annual or perennial herbs with entire opposite leaves

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
    • ]. 

    Bracts persistent, finally ± deflexed; bracteoles and perianth falling with the fruit

  • Provided by: [C].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
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    Distribution

    Of this rather weedy genus of some twenty species, most of the species are centered in Africa and the Orient.

  • Provided by: [E].Flora de Panama
    • Source: [
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     Information From

    Amaranthaceae
    World Flora Online Data. 2017.
    • A CC0 1.0 Universal (CC0 1.0).
    Flora de Nicaragua
    http://www.tropicos.org/projectwebportal.aspx?projectid=7&pagename=Home&langid=66
    WD Stevens, CU Ulloa, A Pool and OM Montiel. Missouri Botanical Garden Press, 2001
    • B Missouri Botanical Garden
    Plants Of the World Online Portal - FTEA
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • C
    Plants Of the World Online Portal - FZ
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • D
    Flora de Panama
    http://www.tropicos.org/Project/PAC
    Robert E. Woodson, Jr. and Robert W. Schery Annals of the Missouri Botanical Garden Vol. 67, No. 4 (1980), pp. ii-xxxiii
    • E Missouri Botanical Garden
    Flora Of CHina @ efloras.org
    http://www.efloras.org/flora_page.aspx?flora_id=2
    'Flora of China @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=2 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
    • F Missouri Botanical Garden
    World Flora Online consortium
    http://www.worldfloraonline.org/organisation/WFO
    World Flora Online Data. 2018.
    • G CC0 1.0 Universal (CC0 1.0).