Plants medium-sized, green, yellowish, or rarely brownish. Stems pinnate or irregularly branched; hyalodermis absent, central strand present; paraphyllia few to many, rarely absent, ; rhizoids or rhizoid initials on stem or abaxial costa insertion, often forming tomentum, usually strongly branched, smooth; axillary hair distal cells 1 or 2, hyaline. Stem leaves not recurved or squarrose, straight or falcate, narrowly to broadly triangular or rounded-triangular, sometimes ovate, not plicate, longer than 1 mm; base decurrent; margins plane or near base slightly recurved, denticulate or serrulate almost throughout, limbidia absent; apex acuminate, ; costa strong, single, usually percurrent or excurrent, sometimes ending well before apex; alar cells differentiated, strongly inflated, hyaline, region well defined, transversely triangular; medial laminal cells elongate-hexagonal, short-rectangular, rectangular, short-linear, or rarely linear, ; marginal cells 1-stratose. Sexual condition dioicous. Capsule horizontal, cylindric, curved; peristome perfect; exostome margins slightly dentate distally; endostome cilia 2 or 3, well developed, nodose. Spores 14-21(-25) µm.