Herbs annual. Leaves alternate, margin lobed; secondary veins ending in sinuses. Inflorescences generally extra-axillary on leafy twigs or in forks of branches, sometimes glomerate. Flowers sessile or on short pedicels, 4-merous. Calyx 4-lobed. Corolla throat glabrous, naked or with 4 scaly appendages; lobes spreading, overlapping in bud. Ovary ovoid, somewhat 4-lobed, 2-loculed and with 2 ovules per locule, or falsely 4-loculed and with 1 ovule per locule; ovules anatropous. Styles 2, terminal, divided or united to middle; stigmas 2, usually divided. Fruit fleshy or dry, fastigiate or semiglobose, endocarp bony, divided into 4 1-seeded achenelike mericarps; mericarps ± united ventrally or joined to a central extension of receptacle. Seeds with little or no endosperm or not; embryo straight or curved.
Ovary 2-locular, each locule with 2 ovules or with 4, 1-ovuled locules; styles 2, terminal, partly joined at the base; stigmas scarcely differentiated
Fruits dry, ovoid-conic, 4-lobed, eventually dividing into 4, 1-seeded nutlets, joined amongst themselves or attached to a central prolongation of the receptacle
Corolla small; tube cylindrical, the throat naked; lobes spreading
Stamens 4, inserted in the tube, included
Flowers solitary, extra- axillary, 4-merous
Leaves alternate, small, numerous, crenate, subsessile or petiolate
Procumbent herb with slender branched stems, often with adventitious roots
Leaves alternate, usually numerous, crenate or lobate, subsessile or petiolate.
Fruit dry or slightly succulent, ± ovoid-pyramidal, 4-lobed, usually separating into 4, 1-seeded nutlets.
Herbs with usually procumbent branched stems.
Flowers 4-merous, solitary, extra-axillary, the upper ones in leafy spicate branches.
Corolla actinomorphic, small, naked.
Stamens inserted below the middle of the corolla tube, included.
Styles 2, slightly united below, terminal; stigmas punctate.