Plants medium-sized to very large, green, pale green, yellowish, brownish, or sometimes pale pink. Stems ± densely radially branched; hyalodermis absent, central strand present; outer pseudoparaphyllia broad, apex rounded to acuminate or cleft; rhizoids or rhizoid initials at various points on leaves, or on scattered points or in rows on stem; axillary hairs , distal cells 2-8(-10), hyaline. Stem leaves ± broadly ovate to broadly rounded-triangular, abruptly narrowed to apex, straight, concave or strongly so, not plicate; base broadly decurrent; margins entire or occasionally slightly sinuate; apex rounded or obtuse; costa single, ending just before apex, or branched to 2-fid and ending 1/2-9/10 leaf length; alar cells differentiated, rectangular to long-rectangular, strongly inflated, hyaline, walls thin, region distinctly or indistinctly delimited, transversely triangular to broadly ovate; medial laminal cell walls incrassate or thin, porose or not. Sexual condition dioicous or autoicous; inner perichaetial leaves not plicate; vaginula naked. Capsule with annulus not separating; exostome external surface reticulate basally, margins dentate distally. Spores 13-31 µm.