Plants small to medium-sized, in moderately loose to dense tufts, green or yellowish to brownish. Stems creeping to ascending, , unevenly foliate, julaceous or not, irregularly to regularly pinnate, branches moderately densely terete- to complanate-foliate, sometimes julaceous; central strand present; pseudoparaphyllia acute; axillary hairs of 3 or 4 cells. Stem leaves erect, patent, or rigidly spreading, imbricate-appressed, ovate, ovate-triangular, or lanceolate, moderately to strongly concave, not or slightly plicate; base decurrent; margins serrulate proximally, serrate distally; apex gradually tapered, acute, acuminate, truncate, apiculate, or cucullate; costa to 40-80% leaf length, broad throughout, terminal abaxial spine present or absent; alar cells subquadrate to short-rectangular, large; laminal cells elongate-flexuose, walls moderately to strongly thick, prorate; basal cells shorter, wider. Branch leaves smaller, narrower; apex acute to acuminate (sharper than stem leaves); costal abaxial surface more strongly serrate; . Sexual condition dioicous; perichaetial leaf acumen reflexed. Seta brownish orange to red-brown, rough. Capsule inclined to horizontal, brownish orange to red-brown, cylindric, not or slightly curved; annulus separating by fragments; operculum long-conic, broadly rostrate; peristome xerocastique, perfect. Calyptra naked. Spores 13-18 µm.