Endosperm dorsally compressed, deeply 4-sulcate dorsally with the vittae in the bottom of the grooves, commissural face only slightly excavate.
Fruit glabrous, slightly dorsally compressed, narrowly elliptic-fusiform, with a broad commissure; primary ribs broad and very obtuse, overlapping the valleculae, which are indicated only by narrow slits, the commissural vittae similarly overlain; vittae solitary in each of the valleculae, 2 on the commissural face; stylopodia conical, large; carpophore divided almost to the base into 2 slender segments
Leaves pinnate to sub-bipinnate or trisect with sharply toothed, acuminate leaflets
Petals greenish, emarginate above with an incurved apical lobule
Perennial with a woody rootstock
Flowers polygamous, the inner of each partial umbel ♂, the outer hermaphrodite
Calyx-teeth well-defined, conspicuous