Rhizome 2-6 mm in diam. (without scales), not white waxy. Scales brown (often grayish), without pale border, broad, ovate to oblong-subdeltoid with rounded to acute apex, appressed to rhizome, usually crisped, margins recurved, basifixed with cordate base and much overlapping lobes, 2-5 mm, without multiseptate hairs, lacking marginal setae or teeth or those rare. Stipe pale, adaxially grooved, 10-20 cm, glabrous or with few scales (sometimes with more scales); lamina compound, tripinnate or quadripinnate, deltoid and broadest toward base, to elongate, often narrowing toward base, 12-50 × 7-40 cm, glabrous, not or slightly dimorphic. Longest petiolules 3-20 mm; pinnae deltoid or linear-triangular; longest pinnae 5-21 × 3-12 cm; pinnules of at least larger pinnae anadromous, linear-oblong or narrowly ovate; longest pinnules 25-70 × 10-35 mm; ultimate pinnae linear-oblong, lobed almost to midrib (each lobe bilobed again); ultimate segments or lobes obtuse or acute without a tooth, 0.5-3 × 0.6-1 mm. Rachises and costae glabrous. Veins in sterile ultimate lobes frequently simple, not reaching margin; false veins not present. Sori separate, frequently single on a segment, at forking point of veins or at bending point of a vein; indusium reniform or semicircular, attached at narrow, cordate base only, wider than long, 0.5-0.8 × 0.5-1 mm.