Roots not proliferous. Stems short-creeping, unbranched; scales dark brown to black throughout, narrowly linear-deltate, 2--4 × 0.3--0.6 mm, margins entire. Leaves ± dimorphic; fertile leaves taller and more erect than sterile leaves. Petiole reddish brown throughout, lustrous, 1--10 cm, 1/4--1/3 length of blade; indument of dark brown to black, filiform scales at base. Blade lustrous, linear to narrowly oblanceolate, 1-pinnate throughout, 4--50 × 2--5(--7) cm, thin, glabrous, or occasionally sparsely pubescent; base gradually tapered; apex acute, not rooting. Rachis reddish or purplish brown throughout, lustrous, glabrous. Pinnae in 15--45 pairs, oblong to quadrangular; medial pinnae 1--2.5 × 0.3--0.5 cm; base with conspicuous acroscopic and sometimes basiscopic auricle, this overlapping rachis; margins crenate to serrulate, sometimes more deeply incised in robust specimens; apex acute to obtuse. Veins free, evident. Sori 1--12 pairs per pinna, on both basiscopic and acroscopic sides. Spores 64 per sporangium. 2 n = 72.
Terrestrial. Rhizome shortly creeping to suberect, with blackish brown scales. Fronds tufted, strongly erect, pinnate, lamina linear-elliptic to linear-oblanceolate, ± 240 x 26 mm, glabrous adaxially, subglabrous with minute, scattered scales abaxially; pinnae roughly rectangular, somewhat falcate and angled downwards, sessile, with enlarged acroscopic lobe, margins crenate to subentire; rachis black, glossy; stipe blackish brown, glabrous at maturity. Sporangia in linear, indusiate sori borne in 7-18 pairs set at 45° angle to midrib.
Rhizome suberect. Fronds to 24 cm, erect, tufted, stipe initially scaled, lamina linear, 1-pinnate, with up to 50 pinna pairs, pinnae oblong, auriculate acroscopically, margins irregularly crenate-serrate, sparsely scaled beneath. Sori to 14 pairs per pinna, indusium erose.
Rhizome erect to suberect, c. 4 mm in diameter, set with dark brown to black, lanceolate-attenuate, subentire rhizome-scales c. 3 mm long. Fronds tufted, firmly membranous, not gemmiferous; stipe atrocastaneous, nitid, less than V6 lamina length, set at first with hair-like scales; lamina linear, c. 240 x 26 mm, pinnate, lower pinnae decrescent; pinnae in up to 50 pairs, subsessile, set at 90 or more to rhachis; midpinnae linear to oblong or lanceolate, auriculate acroscopically, base cordate, overlapping rhachis, margin somewhat irregularly crenate-serrate, incisions sometimes alternating deeper and shallower, upper surface glabrous, lower surface set with minute pale or dark-tipped scales; venation obscure. Sori up to 18 per pinna, c. 2 mm long; indusium membranous, erose, c. 0.5 mm broad.
Mesic Grassland. Forest margins, among rocks and boulder bases in scrub and open grassland.
|Ebony spleenwort, doradille ébène|