Cystopteris protrusa (Weath.) Blasdell
  • Mem. Torrey Bot. Club
  • Southern bladder fern, lowland brittle fern

This taxon is accepted by Cystopteridaceae
Notes: More details could be found in The Plant List v.1.1. Originally in The Plant List v.1.0

General Information

Stems creeping, not cordlike, internodes long, 0.5--1 cm, with persistent petiole bases, covered with tan to golden hairs, especially toward apex; scales tan to light brown, ovate-lanceolate to lanceolate, radial walls thin, luminae tan, mostly crowded at stem apex. Leaves seasonally dimorphic, clustered 1 to several cm beyond persistent old petiole bases, especially in late spring and early summer, to 45 cm, bearing sori (earliest leaves smaller, sterile, coarsely divided, margins with rounded teeth; subsequent leaves larger, fertile, more finely divided, margins with sharply pointed teeth). Petiole mostly green to straw-colored throughout, shorter than or nearly equaling blade, base sparsely scaly. Blade ovate to elliptic, 1-pinnate-pinnatifid to 2-pinnate, widest at or just below middle, apex broadly acute; rachis and costae lacking gland-tipped hairs or bulblets; axils of pinnae lacking multicellular, gland-tipped hairs. Pinnae usually perpendicular to rachis, not curving toward blade apex, margins dentate to serrate; proximal pinnae pinnatifid to pinnate-pinnatifid, ± equilateral, basiscopic pinnules not enlarged, basal basiscopic pinnules stalked, base truncate to obtuse; distal pinnae deltate to ovate. Veins mostly directed into teeth. Indusia ovate to cup-shaped, lacking gland-tipped hairs. Spores spiny, usually 28--34 µm. 2 n = 84.

  • Provided by: [C].Flora of North America @
    • Source: [
    • 1
    • ]. 

    "Rhizome long-creeping, sparsely scaly but densely villous, the growing point produced 2-4 cm beyond the often more scattered lvs; petiole greenish or stramineous except often at the very base; minor axes of the blade with scattered glands; veins typically running into marginal teeth; lowermost pinnules mostly cuneate at the base, and often short-petiolulate; indusium ca 0.5 mm, usually entire; 2n=84; otherwise much like no. 1 [Cystopteris fragilis (L.) Bernh.]. Abundant in mesic woods; N.Y. and s. Ont. to s. Minn., s. to Ga., Ala., La., and e. Kans. (C. fragilis var. p.)"

  • Provided by: [B].New York Botanical Garden
    • Source: [
    • 2
    • ]. 

    Other Local Names

    Southern bladder fern, lowland brittle fern


     Information From

    World Flora Online Data. 2017.
    • A CC0 1.0 Universal (CC0 1.0).
    New York Botanical Garden
    Descriptions of plants should be attributed to the full citation for each individual article, chapter or book that is the source for each record, which should include the authors of original publication.
    • B Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License
    Flora of North America @
    'Flora of North America @ eFloras (2008). Published on the Internet [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
    • C Flora of North America Association
    World Flora Online consortium
    World Flora Online Data. 2018.
    • D CC0 1.0 Universal (CC0 1.0).