Asplenium trichomanes L.
  • Sp. Pl.
  • Saçakotu


Cite taxon page as 'WFO (2020): Asplenium trichomanes L. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-0001109364. Accessed on: 27 Sep 2020'

General Information

Plants 10-30 cm tall. Rhizome erect, short, scaly; scales narrowly triangular, 3(-4) × ca. 0.5 mm, with opaque, red to dark brown central stripe and paler narrow clathrate borders, entire. Fronds caespitose; stipe shiny castaneous-brown, 2-8 cm, base scaly, upward subglabrous, abaxially semiterete but adaxially grooved, with brown, membranous and subentire narrow wings, texture papery, stipe and rachis usually persisting after shedding of pinnae; lamina linear, 10-25 × 0.9-1.6 cm, base slightly reduced, 1-pinnate, apex acute and 2-4 mm wide; pinnae 20-30 pairs, usually obliquely inserted, sessile, middle pinnae elliptic or ovate to orbicular, 2.5-7.5 × 2-4 mm, base nearly symmetrical, cuneate, margin crenate, apex obtuse; lower pinnae gradually reduced. Veins pinnate, costa obscure, veins obliquely simple or up to 2-forked, basal acroscopic vein usually 2-forked. Fronds papery, green or brown when dry, average stomatal guard cell length 35-42 µm; rachis castaneous, shiny, subglabrous, abaxially terete, adaxially grooved and with a relatively low lateral brown membranous wing. Sori 4-8 per pinna, oval to linear, 1-3.5 mm, usually on acroscopic vein; indusium white or brown after drying, oval to linear, membranous, free margin repand to entire, opening toward costa, persistent. Spores with lophate perispore, average exospore length 27-31 µm. Plants sexual diploid: 2n = 72.

  • Provided by: [B].Flora Of CHina @ efloras.org
    • Roots not proliferous. Stems short-creeping, often branched; scales black throughout or with brown borders, lanceolate, 2--5 × 0.2--0.5 mm, margins entire to denticulate. Leaves monomorphic. Petiole reddish brown or blackish brown throughout, lustrous, 1--4(--7) cm, 1/6--1/4 length of blade; indument absent or of black, linear-lanceolate or filiform scales at base. Blade linear, 1-pinnate, 3--22 × 0.5--1.5 cm, thin, glabrous or sparsely pubescent; base gradually tapered; apex narrowly acute, not rooting. Rachis reddish brown throughout, lustrous, glabrous or nearly so. Pinnae in 15--35 pairs, oblong to oval; medial pinnae 2.5--8 × 2.5--4 mm; base broadly cuneate, with or without low, rounded acroscopic auricle; margins shallowly crenate to serrate or ± entire; apex obtuse. Veins free, evident. Sori 2--4 pairs per pinna, on both basiscopic and acroscopic sides. Spores 64 per sporangium. 2 n = 72, 144.

    • Provided by: [G].Flora of North America @ efloras.org
      • Source: [
      • 1
      • ]. 

      "Rhizome 1.5–2 mm thick, the lanceolate brown scales 1.5–2 mm; lvs numerous, 5–25 cm, the purple-brown petiole to 6 cm; blades uniform, linear, pinnate throughout, to 19 × 1.7 cm, acute to attenuate above, usually not narrowed below, thin, dark green, the thin, flexible rachis almost wholly dark; pinnae 9–20 pairs, mostly opposite, oval to round oblong, oblique with the upper side the larger, but not auriculate, 4–8 × 3–6 mm, mostly crenate, subsessile; 2n=72, 144. On shaded, mostly calcareous rocks; interruptedly circumboreal, and nearly throughout our range, s. to Ga. and Tex."

    • Provided by: [C].New York Botanical Garden
      • Source: [
      • 2
      • ]. 

      Morphology

      Çokyıllık, 5-30 cm boyunda demet görünümlü bitkiler. Rizom dik veya yatık, kısa, pullu; pullar ipliksi veya mızraksı, 4 × 0,5 mm, ortası siyah bantlı, kenarları koyu kahverengi ızgaramsı. Fıront sık demet şeklinde, herdem yeşil; sap ve eksen parlak kestane rengi ilâ siyahımsı mor, taban kısmı pullu, üst kısımlar çıplak, sırtı silindirik, karın kısmı oluklu, oluk kenarları iki paralel kahverengi ilâ turuncu renkli dar kanatlı, genellikle telekler döküldükten sonra tel şeklinde üzerinde kalır, 2-8 cm uzunluğunda; fıront ayası, dış hatları şeritsi, 4-25 × 0,9-1,6 cm, koyu yeşil, kâğıtsı, 1 teleksi, 10-32 çift, telekler sapa eğik veya dik bağlanır, telekler kısa saplı veya sapsız, orta kısımdaki telekler dikdörtgenimsi, eliptik, yumurtamsı ilâ dairesel, 2,5-7,5 × 2-4 mm, tabanı hemen hemen simetrik veya asimetrik kamamsı ilâ kesik, üst kenarları kütdişli, tepesi küt, tüysüz; en alttakiler üsttekilere göre yelpazemsi veya üçgensi yuvarlağımsı; damarlar teleksi, orta damar belirgin değil, damarlar eğik, basit veya iki çatalsı. Sorus herbir telekte 4-8 adet, şeritsi 1-3,5 mm, telek orta damarının her iki tarafında, orta damara eğik, telek kenarına kadar uzanmaz; indizyum beyaz, yumurtamsı ilâ şeritsi, zarsı, serbest kenarı hafif dalgalı ilâ düz, orta damar tarafına doğru açılır, kalıcı, olgunlukta sorusu örtmez. Sporlar kahverengi, 27-37 μm.

    • Provided by: [E].Resimli Türkiye Florası (the Illustrated Flora of Turkey)
      • Rhizome erect, set with dark brown rhizome-scales. Pinnae shortly petiolate, broadly oblong-circular, unequally cuneate base forming an angle of 90°, outer margin crenate to dentate. Lamina narrowly linear, up to 150 x 15 mm.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 3
        • ]. 

        Perennial with erect rhizome. Fronds tufted, to 18 cm, stipe glabrous, lamina linear, pinnate, pinnae to 8 mm long, to 32 pairs, broadly oblong-circular, minutely scaled beneath. Null Sori 2-7 per pinna, indusium erose.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 4
        • ]. 

        Rhizome erect, c. 2 mm in diameter, set with lanceolate, attenuate, minutely serrulate, dark brown rhizome-scales up to 4 mm long, often with paler reddish brown margins. Fronds tufted, firmly membranous, gemmiferous; stipe castaneous, usually less than v6 lamina length, glabrous; lamina narrowly linear, up to 150 x 15 mm, somewhat tapering basally, pinnate; pinnae in up to 32 pairs, broadly oblong-circular, the unequally cuneate base forming an angle of 90o, outer margin crenate to dentate, upper surface glabrous, lower surface set with scattered, minute, pale or dark, 1- or 2-celled scales. Sori 2 to 7 per pinna, c. 1.5 mm long; indusium yellowish membranous, erose, c. 0.3 mm broad.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 5
        • ]. 

        Habitat

        Rock crevices and boulder bases.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 3
        • ]. 

        Cliffs and boulders, 800-1400 m.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 4
        • ]. 

        Rock crevices and boulder bases, 1200-3300 m.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 5
        • ]. 

        Distribution

        Tulbagh to Stellenbosch and S Africa to Eurasia, N America, Australasia.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 4
        • ]. 

        Cape Province, Transkei, Natal, Lesotho, Orange Free State, Transvaal, Zimbabwe, Kenya, Ethiopia, Somalia, Socotra, Algeria, Morocco, Canary Islands, Azores, British Isles; North America as far south as Mexico.

      • Provided by: [D].e-Flora of South Africa
        • Source: [
        • 5
        • ]. 

        Nationally Preferred Names

        NameLanguageCountry
        SaçakotuTurkishTR

        Other Local Names

        NameLanguageCountry
        Maidenhair spleenwort, doradille chevelue

          Bibliography

         Information From

        Aspleniaceae
        World Flora Online Data. 2017.
        • A CC0 1.0 Universal (CC0 1.0).
        Flora Of CHina @ efloras.org
        http://www.efloras.org/flora_page.aspx?flora_id=2
        'Flora of China @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=2 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
        • B Missouri Botanical Garden
        New York Botanical Garden
        Descriptions of plants should be attributed to the full citation for each individual article, chapter or book that is the source for each record, which should include the authors of original publication.
        • C Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License
        e-Flora of South Africa
        e-Flora of South Africa. v1.21. 2018. South African National Biodiversity Institute. http://ipt.sanbi.org.za/iptsanbi/resource?r=flora_descriptions&v=1.21
        • D All Rights Reserved
        Resimli Türkiye Florası (the Illustrated Flora of Turkey)
        http://www.turkiyeflorasi.org.tr
        Güner, A., Kandemir, A., Menemen, Y., Yıldırım, H., Aslan, S., Ekşi, G., Güner, I. ve Çimen, A.Ö. (edlr.) (2018). Resimli Türkiye Florası, Cilt 2. ANG Vakfı Nezahat Gökyiğit Botanik Bahçesi Yayınları. İstanbul.
        • E All Rights Reserved
        • F Resimli Türkiye Florası (the Illustrated Flora of Turkey).
        Flora of North America @ efloras.org
        http://www.efloras.org/flora_page.aspx?flora_id=1
        'Flora of North America @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=1 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
        • G Flora of North America Association
        World Flora Online consortium
        http://www.worldfloraonline.org/organisation/WFO
        World Flora Online Data. 2018.
        • H CC0 1.0 Universal (CC0 1.0).