Rhizome 0.5-3 mm in diam. (without scales), white waxy under scales. Scales brown or red-brown, with pale border from base to apex or not, narrowed evenly toward apex, not or seldom curling backward, peltate, 2.5-7 × 0.3-1.5 mm, with multiseptate hairs at least when young, or with marginal setae at least in distal part. Stipe adaxially grooved, 0.1-18 cm, glabrous or with few scales; lamina compound (pinnate with pinnatilobed to pinnatifid pinnae, or bipinnate to quadripinnate toward base and in middle part), simple (margin pectinate or pinnatifid), trifoliolate (pinnae ± divided), or pinnate toward base, ovate, deltoid and broadest toward base, 0.6-24 × 0.5-14 cm, glabrous, strongly dimorphic or not or slightly dimorphic. Longest petiolules 0-4 mm; pinnae linear-triangular, narrowly ovate, linear, or ovate to deltoid; longest pinnae 1-10 × 0.6-7 cm; pinnules (if present) of at least larger pinnae anadromous, linear-oblong or narrowly ovate; longest pinnules 5-55 × 5-20 mm; ultimate pinnae (if present) lobed almost to midrib or only shallowly lobed; ultimate segments or lobes obtuse or acute without a tooth. Dimorphic plants: lamina of fertile fronds pinnate with strongly dissected pinnae, bipinnate, or tripinnate toward base and in middle part; longest petiolules of fertile fronds 1-7 mm; pinnae deltoid, linear-triangular, or narrowly ovate, 1-8 × 0.3-2.5 cm; pinnules or pinna lobes deltoid or linear-oblong, 2-35 × 1.5-15 mm; ultimate pinnae linear-oblong; ultimate segments of fertile fronds 1-15 × 0.5-2 mm. Rachises and costae glabrous. Veins in sterile ultimate lobes simple, forked, or pinnate, reaching margin; false veins not present. Sori separate, borne several on a segment, or in much-divided fronds frequently single on a segment, at forking point of veins; indusium attached at broad base and hardly or not at sides, semicircular or ± triangular to rhomboid, wider than long, ± as wide as long, 0.3-1 × 0.3-1.3 mm, upper margin not elongated, truncate or slightly rounded, separated from or even with lamina margin; lamina generally extending into a tooth at both sides or only at outside of a sorus, or not extending into teeth beyond a sorus.